Monographs
- Pradeu T. (2019), Philosophy of Immunology. Cambridge University Press, Elements in Philosophy of Biology Series (Open Access).
Pradeu T. (2012), The Limits of the Self: Immunology and Biological Identity. New York: Oxford University Press. Lakatos Award 2015. Now available in paperback (January 2019).
Reviews in: Notre Dame Philosophical Reviews (Alfred I. Tauber); Times Literary Supplement (R. P. Novick); Studies in History and Philosophy of Biological and Biomedical Sciences (Neeraja Sankaran); British Journal for the Philosophy of Science (Maureen O’Malley); Biology and Philosophy (Matt Haber); Philosophy in Review (Alison McConwell).
Carosella E.D., Pradeu T. (2010), L’Identité: La Part de l’Autre. Paris: Odile Jacob.

Edited volumes
Gayon J. & Pradeu T. (eds., 2021), Textes Clés en Philosophie de la Biologie. Tome 1 : Explication biologique, hérédité, développement. Tome 2 : Évolution, environnement, diversité biologique. Paris, Vrin. (Key readings in Philosophy of Biology, translated into French, with topic overviews).
Loison L. & Pradeu T. (eds. 2021), La Biologie au défi de l’histoire : Mélanges offerts à Michel Morange. Paris, Éditions Matériologiques.
Guay A. and Pradeu T. (eds.) (2016) Individuals Across the Sciences. New York: Oxford University Press. Contributors: Stéphane Chauvier; Ruey-Lin Chen; Christina Conroy; Mark Ereshefsky & Makmiller Pedroso; Melinda Fagan; Steven French; David Glick; Peter Godfrey-Smith; Alexandre Guay & Thomas Pradeu; Matt Haber; Décio Krause & Jonas R. Becker Arenhart; James Ladyman; Jonathan E. Lowe; Matteo Morganti; Cédric Paternotte; Paavo Pylkkanen, Basil J. Hiley & Ilkka Pattiniemi; Simon Saunders.




Edition of special issues
- Pradeu T. (ed., 2019), Special issue on Philosophy of CRISPR-Cas, with a target paper by Eugene Koonin, 6 commentaries, and the author’s reply. Biology & Philosophy 34(9). (Done as Associate editor of B&P).
- Ferner A. & Pradeu T. (eds., (2017), Special issue on “Ontologies of Living Beings“, Philosophy, Theory, and Practice in Biology, Volume 9. Editorial by A. Ferner & T. Pradeu. Contributors to this special issue: Stéphane Chauvier, Melinda Fagan, Fridolin Gross & Sara Green, Catherine Kendig & Todd Eckdahl, Rory Madden, John Perry, and Christopher Shields. (Open Access).
- Pradeu T. (ed., 2016), Special issue on biological individuality. Biology & Philosophy 31(6), 2016, 761-930. Contributors: Ellen Clarke, Lynn Chiu & Gérard Eberl, Scott Gilbert & Alfred Tauber, Peter Godfrey-Smith, Matt Haber, Thomas Pradeu, David Queller & Joan Strassmann, Derek Skillings.
- Pradeu T., Kostyrka G., and Dupré J. (eds., 2016), Special issue on “Understanding Viruses: Philosophical Investigations“, Studies in History and Philosophy of Science Part C: Studies in History and Philosophy of Biological and Biomedical Sciences 59 (2016), 57-153. Contributors: Jean-Michel Claverie & Chantal Abergel, John Dupré, Patrick Forterre, Eugene Koonin & Petro Starokadomskyy, Gladys Kostyrka, Pierre-Olivier Méthot, Greg Morgan, Maureen O’Malley, Thomas Pradeu, and Marc van Regenmortel.
- Guest editor, special issue on “The Boundaries of development”, Biological Theory 6:1, 1-88. With the contributions of Scott Gilbert, Lucie Laplane, Jane Maienschein, Alessandro Minelli, Michel Morange, Antonine Nicoglou, Thomas Pradeu, Frédérique Théry, and Michel Vervoort.
Articles
- Torri A, Jaeger J, Pradeu T, Saleh M-C (2022), The origin of RNA interference: Adaptive or neutral evolution? PLOS Biology 20(6):e3001715.
- Bazin T, Chiu L, Pradeu T (2022) Host-Microbiota Co-immunity: An Intimate Relationship That Goes Beyond Protection. Philosophy, Theory, and Practice in Biology 14. (Open access).
- Sepich-Poore GD, Guccione C, Laplane L, Pradeu T, Curtius K, Knight R (2022), Cancer’s second genome: Microbial cancer diagnostics and redefining clonal evolution as a multispecies process. BioEssays 44(5), 2100252.
- Laurent P, Lapoirie J, Leleu D, Levionnois E, Grenier C, Jurado-Mestre B, Lazaro E, Duffau P, Richez C, Seneschal J, Pellegrin J-L, Constans J, Schaeverbeke T, Douchet I, Duluc D, Pradeu T, Chizzolini C, Blanco P, Truchetet M-E, Contin-Bordes, C. (2022), Interleukin-1-β-Activated Microvascular Endothelial Cells Promote DC-SIGN+ Alternative Macrophages Associated with Skin Fibrosis in Systemic Sclerosis. Arthritis & Rheumatology 74(6), 1013-1026.
- Sholl J, Sepich-Poore GD, Knight R, Pradeu T (2022), Redrawing therapeutic boundaries: microbiota and cancer. Trends in Cancer 8(2), 87-97. Featured on the cover:
- Kaminski H, Lemoine M, Pradeu T (2021) Immunological exhaustion: How to make a disparate concept operational? PLOS Pathogens Sep 23;17(9):e1009892. doi: 10.1371/journal.ppat.1009892.
- Pradeu T., Lemoine M., Khelfaoui M., Gingras Y. (2021), Philosophy in Science: Can philosophers of science permeate through science and produce scientific knowledge? The British Journal for Philosophy of Science.
- Laurent P, Allard B, Manicki P, Jolivel V, Levionnois E, Jeljeli M, Henrot P, Izotte J, Leleu D, Groppi A, Seneschal J, Constans J, Chizzolini C, Richez C, Duffau P, Lazaro E, Forcade E, Schaeverbeke T, Pradeu T, Batteux F, Blanco P, Contin-Bordes C, Truchetet ME (2021), TGFβ promotes low IL10-producing ILC2 with profibrotic ability involved in skin fibrosis in systemic sclerosis, Annals of the Rheumatic Diseases 80, 1594-1603.
- Khelfaoui M, Gingras Y, Lemoine M, Pradeu T (2021), The visibility of philosophy of science in the sciences, 1980–2018. Synthese. (PDF of final draft).
- Paludan S.R., Pradeu T., Masters S., Mogensen T. (2020), Constitutive immune mechanisms: mediators of host defence and immune regulation. Nature Reviews Immunology 21, 137–150. (Open access).
- Guay A. & Pradeu T. (2020), Right out of the box: How to situate metaphysics of science in relation to other metaphysical approaches. Synthese, 197, 1847–1866. In special issue on “New Metaphysics of Science” edited by Max Kistler. (PDF of final draft).
- Rondeau E., Larmonier N., Pradeu T. and Bikfalvi A. [co-senior authors] (2019) Philosophy of biology: Characterizing causality in cancer eLIFE 8:e53755. (Open Acess).
- Laplane L., Duluc D., Bikfalvi A, Larmonier N., and Pradeu T. (2019), Beyond the tumour microenvironment. International Journal of Cancer 145(10), 2611-2618. (Open Access).
- Pradeu T. (2019), Philosophy of Biology: Immunology and individuality, eLIFE 8:e47384. (Open Access).
- Laplane L., Mantovani P., Adolphs R., Chang H., Mantovani A., McFall-Ngai M., Rovelli C., Sober E., and Pradeu T. (2019), Why science needs philosophy. Proceedings of the National Academy of the Sciences USA (PNAS) 116 (10) 3948-3952. (Open access).
- Bich L., Pradeu T., and Moreau J-F. (2019), Understanding Multicellularity: The Functional Organization of the Intercellular Space. Frontiers in Physiology 10:1170 (Open Access)
- Pradeu T. (2019), Philosophy of CRISPR-Cas. Biology & Philosophy 34(16). (Open access). (PDF of final draft).
- Pradeu T. & Moreau J-F. (2019), CRISPR-Cas immunity: Beyond nonself and defence. Biology & Philosophy 34(6). (PDF of final draft).
- Laplane L., Duluc D., Larmonier N., Pradeu T. & Bikfalvi A. [co-senior authors] (2018), The Multiple Layers of the Tumor Environment. Trends in Cancer 4(12), 802-809. (PDF of final draft). Featured on the cover:
- Lemoine M. & Pradeu T. (2018), Dissecting the meanings of ‘Physiology’ to assess the vitality of the discipline. Physiology 33(4), 236-245. (Open Access PDF).
- Pradeu T. & Du Pasquier L. (2018), Immunological memory: What’s in a name?Immunological Reviews 283(1) (Special Issue: Immunologic Memory), 7-20. (PDF of final draft).
- Truchetet M-E. & Pradeu T. (2018), Re-thinking our understanding of immunity: Robustness in the tissue reconstruction system. Seminars in Immunology 36, 45-55. (PDF of final draft).
- Eberl G. & Pradeu T. (2018), Towards a General Theory of Immunity? Trends in Immunology 39 (4), 261-263. (PDF of final draft).
- Chiu L., Bazin T. (co-first authors), Truchetet M-E., Schaeverbeke T., Delhaes L. & Pradeu T. (2017), Protective Microbiota: From Localized to Long-Reaching Co-Immunity. Frontiers in Immunology, December 2017, Volume 8, Article 1678. (Open Access).
- Ferner A. & Pradeu T. (2017) Editorial introduction to special issue on “Ontologies of Living Beings”, Philosophy, Theory, and Practice in Biology, Volume 9. (Open Access).
- Laurent P., Jolivel V., Manicki P., Chiu L., Contin-Bordes C., Truchetet M-E, and Pradeu T. (co-senior authors) (2017), Immune-Mediated Repair: A Matter of Plasticity. Frontiers in Immunology, Volume 8, Article 454. (Open Access)
- Pradeu T. (2017) Thirty years of Biology & Philosophy: Philosophy of which biology? Biology & Philosophy32(2), 149–167. (PDF of final draft).
- Moreau J-F., Pradeu T., Grignolio A., Nardini C., Castiglione F., Tieri P., Capri M., Salvioli S., Taupin J-L., Garagnani P. & Franceschi C. (2017), The emerging role of ECM crosslinking in T cell mobility as a hallmark of immunosenescence. Ageing Research Reviews 35, 322–335. (Open Access).
- Pradeu T. (2016), Editorial introduction: The many faces of biological individuality. Special issue on biological individuality (T. Pradeu, ed.), Biology & Philosophy31(6), 761-773. (PDF of final draft).
- Pradeu T. (2016), Organisms or biological individuals? Combining physiological and evolutionary individuality. Special issue on biological individuality (T. Pradeu, ed.), Biology and Philosophy31(6), 797–817. (PDF of final draft).
- Pradeu T. & Vivier E. (2016), The Discontinuity Theory of Immunity. Science Immunology1(1), pp. aag0479, July 2016. (PDF of final draft).
- Pradeu T. (2016), Mutualistic viruses and the heteronomy of life, Studies in History and Philosophy of Biological and Biomedical Sciences 59, 80–88. (Long final draft).
- Pradeu T., Kostyrka G., and Dupré J (2016), Understanding viruses: Philosophical investigations, Studies in History and Philosophy of Biological and Biomedical Sciences 59, 57-63. (PDF of final draft)
- Pradeu T, Laplane L, Prévot K, Hoquet T, Reynaud V, Fusco G, Minelli A, Orgogozo V, and Vervoort M. (2016), Defining ‘development’, Current Topics in Developmental Biology, 117, 171-183, special anniversary issue edited by Paul Wassarman. (PDF of final draft)
- Pradeu T. (2014), Galatea of the Microbes. The Philosophers’ Magazine 67 (4th quarter 2014), 89-95.
- Pradeu T., Jaeger S., Vivier E. (2013,) The Speed of Change: Towards a Discontinuity Theory of Immunity? Nature Reviews Immunology 13, 764–769. (PDF of final draft). Featured on the cover:
- Pradeu T., Cooper E.L. (2012), The Danger Theory: Twenty Years Later. Frontiers in Immunology 3: 287 (Open Access paper).
- Pradeu T. (2011), A Mixed Self: The Role of Symbiosis in Development. Biological Theory 6(1): 80-88. (PDF of final draft).
- Pradeu T., Laplane L., Morange M., Nicoglou A., Vervoort M. (2011), Editorial: The Boundaries of Development. Biological Theory 6(1): 1-3. (Open Access).
- Pradeu T. (2010), The Organism in Developmental Systems Theory. Biological Theory 5(3): 216-222. (PDF of final draft).
- Barberousse A., Merlin F., Pradeu, T. (2010), Introduction: Reassessing Developmental Systems Theory. Biological Theory 5(3): 199-201. (PDF of final draft).
- Pradeu T. (2010), What is an organism? An immunological answer. History and Philosophy of the Life Sciences 32: 247-268.
- Pradeu T. (2009), Immune System: “Big Bang” in Question. Science 325: 393.
- Pradeu T. & Carosella E.D. (2006), On the definition of a criterion of immunogenicity. Proceedings of the National Academy of the Sciences USA (PNAS), 103(47): 17858-61. (Open Access paper).
- Carosella E.D., Pradeu T. (2006), Transplantation and Identity: A Dangerous Split. The Lancet 368(9531): 183-184. (PDF of final draft).
- Pradeu T., Carosella E.D. (2006), The Self Model and the Conception of Biological Identity in Immunology. Biology and Philosophy 21(2): 235-252. (PDF of final draft).
Articles in French
- Braillard P-A., Guay A., Imbert C., Pradeu T. (2011), Une objectivité kaléidoscopique : la construction de l’image scientifique du monde. Philosophie 111: 46-71.
- Pradeu T. (2010), L’apport de la philosophie à la question de l’immunogénicité. La Presse Médicale 39(7-8): 747-752.Complément au précédent : Pradeu T. (2010) Réponse à M. Claude Carnaud. La Presse Médicale.Pradeu T. (2009), La mosaïque du soi : les chimères en immunologie. Bulletin de la Société d’Histoire et d’Epistémologie des Sciences de la Vie16(1): 19-27.Pradeu T. (2008), Ces Aliens qui vivent en nous : du parasitisme au piratage génétique. Critique 733-734: 496-509.Pradeu T., Carosella E.D. (2007), Quand et pourquoi une réponse immunitaire est-elle déclenchée? Bulletin de la Société Française d’Immunologie 115: 7.Pradeu T. (2005), Les incertitudes du soi et la question du bon modèle théorique en immunologie. Médecine/Sciences21(10): 872-875.Pradeu T., Carosella E.D. (2005), Tambalea el dogma Propio/No-propio de la immunologia (Questioning the Self/Non-Self Immunology Dogma). Medicina (Buenos Aires)65(6): 558.Pradeu T., Carosella E.D. (2004), Analyse critique du modèle immunologique du soi et du non-soi et de ses fondements métaphysiques implicites. Comptes rendus de l’Académie des sciences Biologies327: 481-492.
Book chapters
- Pradeu T. (2020), Entrée “Organisme”, in Gayon J. (ed.), Dictionnaire encyclopédique de l’identité, Gallimard.Pradeu T. (2020), Entrée “Immunité”, in Gayon J. (ed.), Dictionnaire encyclopédique de l’identité, Gallimard.Pradeu T. (2018), Philosophy of Biology, in. A. Barberousse, D. Bonnay & M. Cozic (eds.), The Philosophy of Science: A Companion. Oxford University Press. (PDF of final draft).Pradeu T. (2018), Jean Gayon, l’histoire, la philosophie et la biologie : Une synthèse moderne, in F. Merlin & Ph. Huneman (eds.), Philosophie, histoire, biologie. Mélanges offerts à Jean Gayon. Paris, Editions Matériologiques. (PDF of final draft).Pradeu T. (2018), Genidentity and Biological Processes, in Nicholson, D. J. & Dupré, J. (eds.). Everything Flows: Towards a Processual Philosophy of Biology. Oxford: Oxford University Press, pp. 96-112. Open access to the whole book in PDF file.
- Guay A. and Pradeu T. (2016), Introduction: Progressive Steps Towards a Unified Conception of Individuality Across the Sciences, in Guay and Pradeu, eds., Individuals Across the Sciences, pp. 1-21.
- Guay A. and Pradeu T. (2016), To be continued: the genidentity of physical and biological processes, in Guay and Pradeu, eds., Individuals Across the Sciences, pp. 317-347.
- Minelli A. & Pradeu T. (2014), Theories of development in biology – problems and perspectives, in A. Minelli and T. Pradeu, eds., Towards a Theory of Development. Oxford: Oxford University Press, pp. 1-14.
- Pradeu T. (2014), Regenerating Theories in Developmental Biology, in A. Minelli and T. Pradeu (eds.) Towards a Theory of Development. Oxford: Oxford University Press, pp. 15-32.
- Pradeu T. (2013), Immunity and the emergence of individuality, in F. Bouchard and P. Huneman (eds.) From Groups to Individuals: Perspectives on Biological Associations and Emerging Individuality. Cambridge, MA: MIT Press, pp. 77-96.
- Pradeu T. (2013), Critique des notions de soi et de non-soi en immunologie, in J-J. Kupiec (ed.) La Vie, et alors? Paris: Belin, pp. 177-189.
- Pradeu T. (2011), Chapitre “Philosophie de la biologie”, in A. Barberousse, D. Bonnay, M. Cozic (eds.) Précis de Philosophie des Sciences. Paris, Vuibert, 378-403. (PDF of final draft).
- Pradeu T. (2010), Peut-on attribuer une fonction au système immunitaire?, in J. Gayon & A. de Ricqlès (eds.) Les Fonctions : des organismes aux artefacts. Paris: Presses Universitaires de France, pp. 261-275.
- Pradeu T. (2009), Darwinisme, évolution et immunologie, in T. Heamns, P. Huneman, G. Lecointre, M. Silberstein (eds.), Les Mondes darwiniens: L’évolution de l’évolution. Paris: Syllepses, pp. 525-542.Site web: http://www.lesmondesdarwiniens.org/
Nouvelle édition parue en 2011 aux éditions Matériologiques: http://www.materiologiques.com/Les-mondes-darwiniens-L-evolution
- Pradeu T. (2008), Qu’est-ce qu’un individu biologique?, in P. Ludwig, T. Pradeu (eds.), L’individu : Perspectives contemporaines. Paris : Vrin, pp. 97-125. (PDF)
- Pradeu T. (2007), L’immunité et l’interactionnisme biologique, in Thierry Martin (dir.), Le Tout et les parties dans les systèmes naturels. Paris: Vuibert, coll. “Philosophie des sciences”, pp. 99-106.
- Carosella E.D., Pradeu T. (2006), Introduction, in Carosella E.D., Pradeu T., Saint-Sernin B., Debru C. (eds.) L’identité? Soi, non-soi, individu et personne. Paris: PUF, pp. 3-5.Pradeu T., Carosella E.D. (2006), L’identité immunologique : soi ou continuité?, in Carosella E.D., Pradeu T., Saint-Sernin B., Debru C. (eds.) L’identité? Soi, non-soi, individu et personne. Paris: PUF, pp. 47-58.
Book reviews
- Pradeu T. (2016), Toolbox Murders: Putting Genes in their Epigenetic and Ecological Contexts: Review of Griffiths and Stotz, Genetics and Philosophy: An Introduction (Cambridge University Press, 2013). Biology & Philosophy, 31(1), 125-142. Preprint (longer version of the same paper): here.Pradeu T. (2011), Review of P. Godfrey-Smith’s Darwinian populations and natural selection, (Oxford University Press, 2009), Mind 120(479): 863-870.Pradeu T. (2011), What philosophy of biology should be. Review of A. Rosenberg and D. McShea, Philosophy of Biology. A Contemporary Introduction, Routledge, 2008. Biology and Philosophy26: 119-127.
Outreach activities
- Pradeu T. (2021), Interview for “New Books in Philosophy” about my book Philosophy of Immunology. (February 2021).Pradeu T. (2021), Participation au dossier « Bactéries : elles font aussi ce que nous sommes », Science & Vie N°1240, p. 68-87. (Janvier 2021).Pradeu T. (2020), « L’immunité collective doit être liée à la vaccination », Interview pour Le Point.Pradeu T. (2020), Participation à une « enquête sur l’immunité collective », Philosophie Magazine. (Mars 2020).Pradeu T. (2020), « Cette crise est une leçon d’humilité et d’altruisme », Interview pour La Vie.Pradeu T. (2017), On peut identifier un individu grâce aux caractéristiques uniques de son microbiote. Interview dans In Vivo Magazine (Revue du CHUV), Numéro “Les bactéries, nos meilleures ennemies”, n°13, Décembre 2017, pp. 28-29.
- Participation au livre Etonnant vivant : Découvertes et promesses du XXIe siècle, dir. Catherine Jessus, Paris, CNRS Editions. (Contribution au Chapitre 1 “Qu’est-ce que le vivant?”).
- Pradeu T. (2016), Vers une nouvelle vision de l’immunité. Pour la Science, numéro spécial “Le nouveau monde des microbes”, N°469, Novembre 2016, pp. 62-63.
- Pradeu T. (2015), « Philosophie de la biologie », Dossier « Les nouveaux enjeux de la philosophie », Sciences Humaines, avril 2015.Interview, Le Monde, 18 Nov 2013 : “La philosophie nourrit les sciences et se nourrit d’elles
Other
- Pradeu T. (2009), Obituary: Marie-Claude Lorne. Biology and Philosophy 24(3): 281-282.